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Richard H Shukle


  • Adjunct Professor of Entomology
  • Adjunct Professor
  • Associate Professor / USDA-ARS Entomologist / Hessian Fly Genetics

Selected Publications

Stuart, J. J., Harris, M. O., Chen, M. S., & Shukle, R. H. (2012). Gall midges (Hessian flies) as plant pathogens. Annual Review of Phytopathology, 50, 339-357.

Johnson, A. J., Weintraub, P. G. H., Katoch, R., Cambron, S. E., Schemerhorn, B. J., & Shukle, R. H. (2012). Biological and molecular characterization of Hessian fly (Diptera: Cecidomyiidae) from Israel. Bulletin of Entomological Research, 102, 632-643.

Shukle, R. H., Subramanyam, S., & Williams, C. E. (2012). Effects of antinutrient proteins on Hessian fly (Diptera: Cecidomyiidae) larvae. Journal of Insect Physiology, 58, 41-48.

Johnson, A. J., Morton, P. K., Schemerhorn, B. J., & Shukle, R. H. (2011). Use of a nuclear marker to assess population structure in Hessian fly (Mayetiola destructor). Annals of the Entomological Society of America, 104, 666-674.

Mittapalli, O., Rivera-Vega, L., Bhandary, B., Bautista, M. A., Mamidala, P., Michel, A. P., . . . Mian, M. A. R. (2011). Cloning and characterization of mariner-like elements in soybean aphid, Aphis glycines Matsumura. Bulletin of Entomological Research, 101, 697-704.

Williams, C. E., Nemacheck, J. A., Shukle, J. T., Subramanyam, S., Saltzmann, K. D., & Shukle, R. H. (2011). Induced epidermal permeability modulates resistance and susceptibility of wheat seedlings to herbivory by Hessian fly larvae. Journal of Experimental Biology, 62, 4521-4531.

Chen, M. S., Liu, X. M., Yang, Z., Zhao, H., Shukle, R. H., Stuart, J. J., & Hulbert, S. (2010). Unconventional conservation among genes encoding small secreted salivary gland proteins from a gall midge. BMC Evolutionary Biology, 10, 296. Retrieved from

Behura, S. K., Shukle, R. H., & Stuart, J. J. (2010). Assessment of structural variation and molecular mapping of insertion sites of Desmar-like elements in the Hessian fly genome. Insect Molecular Biology, 19(6), 707-715. Retrieved from

Cambron, S. E., Buntin, G. D., Weisz, R., Holland, J. D., Flanders, K. L., Schemerhorn, B. J., & Shukle, R. H. (2010). Virulence in Hessian fly (Diptera: Cecidomyiidae) field collections from the southeastern United States to twenty-one resistance genes in wheat. Journal of Economic Entomology, 103(6), 2229-2235. Retrieved from

Shukle, R. H., Subramanyam, S., Saltzmann, K. A., & Williams, C. E. (2010). Ultrastructural changes in the midguts of Hessian fly larvae feeding on resistant wheat. Journal of Insect Physiology, 56(7), 754-760. Retrieved from